In the beginning of modern plant biology, plant biologists followed a simpl
e model for their science. This model included important branches of plant
biology known then. Of course, plants had to be identified and classified f
irst. Thus, there was much work on taxonomy, genetics, and physiology. Ecol
ogy and evolution were approached implicitly, rather than explicitly, throu
gh paleobotany, taxonomy, morphology, and historical geography. However, th
e burgeoning explosion of knowledge and great advances in molecular biology
, e.g., to the extent that genes for specific traits can be added (or delet
ed) at will, have created a revolution in the study of plants. Genomics in
agriculture has made it possible to address many important issues in crop p
roduction by the identification and manipulation of genes in crop plants. T
he current model of plant study differs from the previous one in that it pl
aces greater emphasis on developmental controls and on evolution by differe
ntial fitness. In a rapidly changing environment, the current model also ex
plicitly considers the phenotypic variation among individuals on which sele
ction operates. These are calls for the unity of science. In fact, the prop
onents of "Complexity Theory" think there are common algorithms describing
all levels of organization, from atoms all the way to the structure of the
universe, and that when these are discovered, the issue of scaling will be
greatly simplified! Plant biology must seriously contribute to, among other
things, meeting the nutritional needs of the human population. This challe
nge constitutes a key part of the backdrop against which future evolution w
ill occur. Genetic engineering technologies are and will continue to be an
important component of agriculture; however, we must consider the evolution
ary implications of these new technologies. Meeting these demands requires
drastic changes in the undergraduate curriculum. Students of biology should
be trained in molecular, cellular, organismal, and ecosystem biology, incl
uding all living organisms.