Near the end of the Late Ordovician, in the first of five mass extinctions
in the Phanerozoic, about 85% of marine species died. The cause was a brief
glacial interval that produced two pulses of extinction. The first pulse w
as at the beginning of the glaciation, when sea-level decline drained epico
ntinental seaways, produced a harsh climate in low and mid-latitudes, and i
nitiated active, deep-oceanic currents that aerated the deep oceans and bro
ught nutrients and possibly toxic material up from oceanic depths. Followin
g that initial pulse of extinction, surviving faunas adapted to the new eco
logic setting. The glaciation ended suddenly, and as sea level rose, the cl
imate moderated, and oceanic circulation stagnated, another pulse of extinc
tion occurred. The second extinction marked the end of a long interval of e
cologic stasis tan Ecologic-Evolutionary Unit). Recovery from the event too
k several million years, but the resulting fauna had ecologic patterns simi
lar to the fauna that had become extinct. Other extinction events that elim
inated similar or even smaller percentages of species had greater long-term
ecologic effects.