Zc. Jewell et al., Censusing and monitoring black rhino (Diceros bicornis) using an objectivespoor (footprint) identification technique, J ZOOL, 254, 2001, pp. 1-16
An objective, non-invasive technique was developed for identifying individu
al black rhino from their footprints (spoor). Digital images were taken of
left hind spoor from tracks (spoor pathways) of 15 known black rhino in Hwa
nge National Park, Zimbabwe. Thirteen landmark points were manually placed
on the spoor image and from them, using customized software, a total of 77
measurements (lengths and angles) were generated. These were subjected to d
iscriminant and canonical analyses. Discriminant analysis of spoor measurem
ents from all 15 known animals, employing the 30 measurements with the high
est F-ratio values, gave very close agreement between assigned and predicte
d classification of spoor. For individual spoor, the accuracy of being assi
gned to the correct group varied from 87% to 95%. For individual tracks, th
e accuracy level was 88%. Canonical analyses were based on the centroid plo
t method, which does not require pre-assigned grouping of spoor or tracks.
The first two canonical variables were used to generate a centroid plot wit
h 95% confidence ellipses in the test space. The presence or absence of ove
rlap between the ellipses of track pairs allowed the classification of the
tracks. Using a new 'reference centroid value' technique, the level of accu
racy was high (94%) when individual tracks were compared against whole sets
(total number of spoor for each rhino) but low (35%) when tracks were comp
ared against each other. Since tracks with fewer spoor were more likely to
be misclassified, track sizes were then artificially increased by summing s
maller tracks for the same rhino. The modified tracks in a pairwise compari
son gave an accuracy of 93%. The advantages, limitations and practical appl
ications of the spoor identification technique are discussed in relation to
censusing and monitoring black rhino populations.