In all terrestrial and aquatic plant species the primary cell wall is a dyn
amic structure, adjusted to fulfil a diversity of functions. However a univ
ersal property is its considerable mechanical and tensile strength, whilst
being flexible enough to accommodate turgor and allow for cell elongation.
The wall is a composite material consisting of a framework of cellulose mic
rofibrils embedded in a matrix of non-cellulosic polysaccharides, interlace
d with structural proteins and pectic polymers. The assembly and modificati
on of these polymers within the growing cell wall has, until recently, been
poorly understood. Advances in cytological and genetic techniques have thr
own light on these processes and have led to the discovery of a number of w
all-modifying enzymes which, either directly or indirectly, play a role in
the molecular basis of cell wall expansion.