Exotic annual weeds on western U.S. rangelands owe at least part of th
eir invasion success to prolific seed production. Reproductive output
is determined largely by timing of flowering in relation to soil moist
ure, light, and competing vegetation. This study investigated the infl
uence of temperature (including vernalization), photoperiod, and intra
specific competition on reproductive phenology in three populations (t
wo varieties, typica and brachypappa) of common crupina. Monthly plant
ings at four densities were grown in field plots through two consecuti
ve cycles, and the dare each plant reached bud and flowering stage was
recorded. Development of greenhouse-grown plants from the same popula
tions was compared with those grown outside for 4, 6, 8, and 10 wk of
cold induction. Reproductive development in common crupina was influen
ced by all factors investigated: temperature, photoperiod, density, an
d vernalization. Conditions fostering the shortest thermal time from e
mergence to floral induction were a period of cold temperatures/short
photoperiod as seedlings followed by warming temperatures and lengthen
ing photoperiod. Both varieties of common crupina responded quantitati
vely to vernalization, but lack of vernalization caused a greater dela
y of flowering in var. brachypappa than in var. typica. Vernalization
contributed to convergence of spring flowering among the August to Dec
ember plantings, but earlier emergence and lower plant density greatly
increased final plant size and fecundity. Lack of vernalization, high
temperatures, and shortening day length delayed flowering of spring/s
ummer emergents, increasing the probability of succumbing to drought b
efore producing seed.