A new temnospondyl amphibian find from the Early Triassic Lystrosaurus Asse
mblage Zone of the Beaufort Group (Karoo Basin) represents the first undoub
ted occurrence of a long-snouted trematosaurid in South Africa. Its discove
ry confirms the presence of trematosaurids In the Lystrosaurus Assemblage Z
one and brings to three the number of diagnostic trematosaurid taxa in the
Beaufort Group, which were previously known only from short-snouted taxa, T
he presence of a derived, long-snouted trematosaurid in basal Triassic rock
s of the Lystrosaurus Assemblage Zone, and that of apparently more primitiv
e short-snouted trematosaurids in the overlying Cynognathus Assemblage Zone
, follows the trend seen elsewhere in Pangaea in which the highly specializ
ed long-snouted trematosaurid taxa appear earlier in the fossil record.