Much research has recently been dedicated to applying genetic algorith
ms to populations of neural networks. However, while in real organisms
the inherited genotype maps in complex ways into the resulting phenot
ype, in most of this research the development process that creates the
individual phenotype is ignored. In this paper we present a model of
neural development which includes cell division and cell migration in
addition to axonal growth and branching. This reflects, in a very simp
lified way, what happens in the ontogeny of real organisms. The develo
pment process of our artificial organisms shows successive phases of f
unctional differentiation and specialization. In addition, we find tha
t mutations that affect different phases of development have very diff
erent evolutionary consequences. A single change in the early stages o
f cell division and migration can have huge effects on the phenotype,
while changes in later stages usually have a less dramatic impact. Som
etimes, changes that affect the first developmental stages may be reta
ined, producing sudden changes in evolutionary history.