In 1787, the U.S. Congress authorized the sale of the "Ohio Company Purchas
e", ca. 5000 km(2) in Appalachian Ohio. The land was surveyed using a towns
hip and range system shortly thereafter. Data on > 5600 witness trees were
transcribed from the survey records, and witness tree locations were plotte
d on a digital map. This information was used to evaluate presettlement for
est composition and structure and to investigate vegetation-site relationsh
ips before widespread alteration of the forests had taken place. Presettlem
ent conditions were compared with present conditions using forest inventory
and analysis (FIA) data. Two hundred years ago, the forests of southeaster
n Ohio were dominated by large individuals of Quercus alba L., Carya Nutt.
spp., Quercus velutina Lam., and Fagus grandifolia Ehrh. These four taxa ac
counted for 74% of all witness trees. Although almost 70% of the region is
forested today, the second-growth forest has witnessed a decrease in Quercu
s and Carya and an increase in Acer saccharum Marsh., Acer rubrum L., and m
any early successional species in smaller size classes. Despite the signifi
cant shift in forest composition and structure, species in general seem to
be occupying similar positions in the present-day landscape compared with t
he presettlement forest; topographic variables most strongly control specie
s occurrence in this landscape.