Phylogenetic relationships of the intertidal genus Halorhadinus Sawada andkey to the genera of the Liparocephalini (Coleoptera : Staphylinidae : Aleocharinae)

Halorhadinus Sawada is redescribed and illustrations of diagnostic features are presented. Median lobes and parameres of H. aequalis Sawada and H. ina equalis Sawada are described for the first time. A key is provided for sepa ration of the known species of Halorhadinus. Based on contiguous mesocoxal cavities, galea with several setae only on mesal surface and apex with seta e, Halorhadinus is hypothesized to be a member of the tribe Liparocephalini . A revised cladistic analysis of the Liparocephalini based on 50 adult cha racters suggests that Halorhadinus belong to the Liparocephalini with the f ollowing patterns of generic relationships (outgroup (Salinamexus Moore & L egner (Halorhadinus Sawada (Amblopusa Casey (Paramblopusa Ahn & Ashe (Diaul ota Casey + Liparocephalus Maklin)))))). Monophyly of the genus Halorhadinu s is supported by three synapomorphies (mentum with anterior margin deeply sinuate and antero-lateral parts triangular; ligula round and very long, al most reaching to half of palpi; spatulate setae mostly located on basal par t of palpi). A key to the genera of the Liparocephalini is given.