Yf. Hsu et Wi. Chou, Araragi panda sp n., an intermediate taxon in the phylogeny of Theclini (Lepidoptera : Lycaenidae : Theclinae), IN SYST EVO, 32(2), 2001, pp. 155-168
The current placement of the genus Araragi in the Chaetoprocta lineage has
been problematic because Araragi does not possess a bilobed uncus, a synapo
morphy shared by all the other genera of the lineage. The discovery of Arar
agi panda, a new species with features of Araragi and a bilobed uncus, just
ifies this systematic treatment. Phylogenetic analysis identifies Chaetopro
cta as the sister of Araragi, and demonstrates that colonization of walnuts
as larval hosts occurred only once in Theclini evolution, contradicting pr
eviously proposed hypotheses.