On the basis of representative collections made from seven measured section
s in western Hubei and Hunan, four early Llanvirn and five late Llanvirn-ea
rly Caradoc trilobite biofacies are distinguished along a bathymetric gradi
ent from shallow to deep outer shelf The diversity and relative abundance o
f mesopelagic cyclopygids gradually increased southwards or seawards. Accor
ding to the facies association exhibited in the region, the upper limit of
living depth for different forms is inferred as follows: 1) more than 70 m
(M. (Microparia)); 2) less than 100 m (M. (Quadratapyge), Pricyclopyge, Asp
idaeglina); 3) at approximately 100 m (Cyclopyge); and 4) more than 100 m.
(Symphysops, Psilacella). Benthic forms were absent in the deep outer-shelf
basin because of stagnant conditions. In shallower sites of the outer shel
f, genera of nileids in the carbonate facies and raphiophorids in the clast
ic facies were relatively abundant but some endemic forms, such as the asap
hid Birmanites and remopleuridid Hexacopyge, became dominant during the lat
e Llanvirn-early Caradoc. Rorringtoniids are mainly associated with deeper
outer-shelf facies. Panderia is common in an early Llanvirn nileid fauna at
Yichongqiao, Cili area, and may well indicate development of shelf carbona
te buildups. A transgressive event is suggested by the sudden appearance of
rich Cyclopyge in the early Caradoc at Jiuxi, Changde.