R. Guidetti et R. Bertolani, An evolutionary line of the Macrobiotinae (Tardigrada, Macrobiotidae): Calcarobiotus and related species, ITAL J ZOOL, 68(3), 2001, pp. 229-233
Three species of Macrobiotus (M. gildae, M. polygonatus and M. eugranulatus
) were re-examined. Based on their synapomorphies with Calcarobiotus, they
are ascribed to that genus. Within Calcarobiouts two phyletic lines are det
ected on the basis of the claw morphology. Therefore a new subgenus is erec
ted: Calcarobiotus (Discrepunguis) subgen. n. constituted by C. (D.) polygo
natus comb. n., and C. (D.) eugranulatus comb. n.; the subgenus Calcarobiot
us (Calcarobiotus) includes C (C) filmeri, C. (C) occultus, C. (C.) imperia
lis and C. (C.) gildae comb. n.