DESCRIPTION OF CRYPTOCERCUS-CLEVELANDI (DICTYOPTERA, CRYPTOCERCIDAE) FROM THE NORTHWESTERN UNITED-STATES, MOLECULAR ANALYSIS OF BACTERIAL SYMBIONTS IN ITS FAT-BODY, AND NOTES ON BIOLOGY, DISTRIBUTION, AND BIOGEOGRAPHY

Previously published studies on distribution, out fauna, glandular sec retions, karyotypes. 12S and 16S mitochondrial ribosomal DNA (rDNA) se quence divergence, and reproductive isolation suggest that at least 2 species have been included in the taxon Cryptocercus punctulatus Scudd er. Here we present morphological and biological studies of the insect s, including 16S rDNA sequence divergence of their bacterial endosymbi onts in the fat body. We conclude that the name C. punctulatus should be restricted to populations from the Appalachian region of the easter n United States. Those in southern Washington, Oregon, and northern Ca lifornia in the northwestern United States are a distinct species here assigned to Cryptocercus clevelandi by G. W. Byers. The genus Cryptoc ercus now consists of 4 species, whose distribution suggests an early Tertiary origin.