Df. Hoese et S. Reader, Revision of the eastern Pacific species of Gobulus (Perciformes : Gobiidae), with description of a new species, REV BIOL TR, 49, 2001, pp. 169-176
Gobulus birdsongi is described as a new species from the Pacific coast of P
anama. The two other known species of Gobulus from the eastern Pacific, Gob
ulus crescentalis and G. hancocki are redescribed. Gobulus birdsongi differ
s from other species in the genus in having more numerous second dorsal and
anal rays. Gobulus hancocki differs from G. crescentalis in having a much
smaller eye. The genus is distinctive in having reversed countershading, wi
th the ventral surface of the body darker than the dorsal surface.