Synechococcus sp. strain PCC 7942 has a second clpB gene that encodes a 97-
kDa protein with novel features. ClpBII is the first ClpB not induced by he
at shock or other stresses; it is instead an essential, constitutive protei
n. ClpBII is unable to complement ClpBI function for acquired thermotoleran
ce. No truncated ClpBII version is normally produced, unlike other bacteria
l forms, while ectopic synthesis of a putative truncated ClpBII dramaticall
y decreased cell viability.