Inclusive fitness theory has been used to explain the overall pattern of al
truistic behavior. However, this theory does not address the proximal cause
s of altruism. The purpose of the present study was to increase the underst
anding of altruism by testing a model that includes both ultimate and proxi
mate causes. In particular, emotional closeness was tested as a mediator of
the effect of genetic relatedness on altruistic behavior. This was accompl
ished by having college students choose which of their family members they
would most likely provide with life-saving assistance. As expected, results
showed that emotional closeness is an important proximal cause of altruism
that partially mediates the effect of genetic relatedness on willingness t
o act altruistically.