Unsteady fluid dynamic farces acting on a two-dimensional wing in sinusoida
l plunging and pitching motions in still water were measured. The measured
fluid dynamic forces were larger than those estimated by quasi-steady analy
sis, where the effect of unsteady separated vortices on the fluid dynamic f
orces is not considered. Vortex capture is a phenomenon related to the larg
er fluid dynamic forces in wing motion that is similar to wing motion of re
al insects. By using the measured fluid dynamic forces, we identified combi
nations of plunging and pitching motions for maximum time-averaged thrust a
nd for maximum efficiency.