GEOGRAPHIC-VARIATION IN RESPONSE OF PINE ENGRAVER, IPS PINI, AND ASSOCIATED SPECIES TO PHEROMONE, LANIERONE

Lanierone strongly synergized the attraction of male and female Ips pi ni (Say) to ipsdienol in New York and Wisconsin. Synergy was only weak ly significant in Montana and British Columbia and not significant in California. Catches of I. pini in ipsdienol-baited traps were increase d 0% (i.e., nonsignificant) to 9942% by lanierone, with the highest in creases in eastern North America. Lanierone had the least effect in Ca lifornia. The effects of lanierone on sex ratios of I. pini in trap ca tches varied significantly between regions. The addition of lanierone to ipsdienol-baited traps resulted in a general increase in male repre sentation at nine of 12 sites. Ips integer (Eichhoff) was attracted to lanierone alone. Ipsdienol reduced the response of I. integer to lani erone. Enoclerus lecontei (Wolcott) (Cleridae) preferred traps baited with the combination of ipsdienol and lanierone. (R)-(-)-Ipsdienol was attractive to E. sphegeus (F.), Thanasimus undatulus (Say) (Cleridae) , and Temnochila chlorodia (Mannerheim) (Trogositidae), while racemic ipsdienol was attractive to E. nigrifrons var. gerhardi Wolc. and Than asimus dubius (F.). Lanierone had no effect on these species.