Jj. Morrone, THE SOUTH-AMERICAN WEEVIL GENUS RHYEPHENES (COLEOPTERA, CURCULIONIDAE, CRYPTORHYNCHINAE), Journal of the New York Entomological Society, 104(1-2), 1996, pp. 1-20
The genus Rhyephenes Schoenherr, endemic to the Central Chilean and Su
bantarctic biogeographic provinces of southern South America, is easil
y recognized by its black body with shape resembling a spider. This ge
nus includes seven species: Rhyephenes clathratus R. Philippi, R. gayi
(Guerin), R. goureaui (Gay & Solier), R. humeralis (Guerin), R. later
alis (Guerin), R. maillei (Gay & Solier), and R. squamiger F. Philippi
(reinstated herein from synonymy with R. gayi). A key, redescriptions
, habitus photographs, and illustrations of the species are provided,
and their geographical distribution is mapped. A cladistic analysis us
ing 21 characters from external morphology and male and female genital
ia produced four cladograms (CI = 0.55, Ri = 0.51, length 43 steps), w
hich after successive weighting were reduced to one cladogram (CI = 0.
82, RI = 0.85, length ill steps). In the cladogram, the following phyl
ogenetic sequence results: (R. squamiger, (R, lateralis, ((R. clathrat
us, R. goureaui), (R. gayi, (R. humeralis, R. maillei))))). Biogeograp
hic patterns exhibited by these species indicate a sequence from centr
al Chile to southern Chile and Argentina.